NATIONAL PARK SERVICE Mountain Goats in Olympic National Park: Biology and Management of an Introduced Species

The U.S. Department of the Interior, National Park Service funded the studies. We thank former Olympic National Park superintendents R. Chandler and R. Contor and current superintendent M. Finnerty for their support. With assistance from former Regional Chief Scientist D. Field and Regional Chief Scientist J. Larson, funds were also provided by the Pacific Northwest and Washington, D.C., offices of the National Park Service. Additional support for several research studies was provided by the University of Washington and the University of Idaho. Assistance in the field was provided by the ranger and naturalist staffs at Olympic National Park. Washington Department of Wildlife personnel aided in goat capture, and the Student Conservation Association provided volunteers to aid fieldwork. J. Foster, D. Jessup, B. Marts, and C. Robbins recommended the medical protocol for captured goats. Numerous seasonal employees aided all aspects of the study, and we thank the following, among others, for their steadfast assistance: K. Bierly, M. Bivin, J. Burger, A. Comulada, M. Dalton, M. Dearstyne, D. Doss, C. Early, J. Enright, D. Frank, R. Fredrickson, G. Hunter, T. Kaye, P. Maurides, P. McElhose, S. Mills, R. Reid, E. Seaman, W. Weins, K. Wilson, and J. Young. J. Mott and other pilots at Eagle Air Helicopters skillfully piloted the mountain goat censuses and capture flights. We thank those who generously shared historic photographs used to document vegetation changes, particularly J. Harter, L. Lack, and the collections of the Klahhane Club and the J. W. Thompson Co.

The editors of the Canadian Journal of Zoology, Northwest Science, and the Journal of Mammalogy gave permission to reproduce Figs. 18, 19, 26, and 31. M. Vavra, Eastern Oregon Agricultural Research Center, Burns, Oregon, and A. Olson, Toutle, Washington, analyzed samples of ungulate diets. G. Caughley, L. L. Eberhardt, and A. R. E. Sinclair provided advice on goat census techniques and interpretations of goat demography. L. Conquest, Center for Quantitative Studies, University of Washington, Seattle, provided statistical advice.

Earlier versions of the material on herbivory benefited greatly from reviews by J. Bailey, L. Bliss, R. del Moral, J. Dennis, R. Fonda, D. Peterson, S. Sharrow, E. Starkey, and G. Wright. Additionally, we appreciate the reviews of Chapters 2 and 3 by C. Whitlock; Chapter 3 by K. Aubry, T. Cox, and M. Greene; and Chapters 1 and 14 by J. Larson and K. Jope. The material in Chapter 3 and Appendix B benefited from reviews by T. Cox, P. Farber, G. Frederick, M. Greene, B. Lane, B. Maurer, M. Moss, H. Smith, and H. Steen. The entire manuscript was read and greatly improved by comments received from J. Agee, C. Hoffman, R. Hoffman, L. Irwin, D. Peterson, E. Starkey, V. Stevens, and three anonymous reviewers.